J. Berman, U. Zorrilla-lópez, G. Farré, C. Zhu, G. Sandmann et al., Nutritionally important carotenoids as consumer products, Phytochemistry Reviews, vol.11, issue.165???188, pp.727-743, 2014.
DOI : 10.1111/j.1467-7652.2012.00740.x

R. Calegari-santos, R. A. Diogo, J. D. Fontana, and T. M. Bonfim, Carotenoid Production by Halophilic Archaea Under Different Culture Conditions, Current Microbiology, vol.197, issue.5, pp.641-651, 2016.
DOI : 10.1128/JB.02523-14

Y. Caro, L. Anamale, M. Fouillaud, P. Laurent, T. Petit et al., Natural hydroxyanthraquinoid pigments as potent food grade colorants: an overview, Natural Products and Bioprospecting, vol.74, issue.5, pp.174-193, 2012.
DOI : 10.1002/bdrb.20043
URL : https://hal.archives-ouvertes.fr/hal-01188117

L. M. Castelblanco-matiz, A. Barbachano-torres, T. Ponce-noyola, A. C. Ramos-valdivia, C. García-rojas et al., Carotenoid production and gene expression in an astaxanthin-overproducing Xanthophyllomyces dendrorhous mutant strain, Archives of Microbiology, vol.37, issue.5, pp.1129-1139, 2015.
DOI : 10.1007/s002940050010

J. Chang, C. Thia, H. Lin, H. Liu, F. Ho et al., Integrating an algal ??-carotene hydroxylase gene into a designed carotenoid-biosynthesis pathway increases carotenoid production in yeast, Bioresource Technology, vol.184, pp.2-8, 2015.
DOI : 10.1016/j.biortech.2014.11.097

Y. Chen, W. Xiao, Y. Wang, H. Liu, X. Li et al., Lycopene overproduction in Saccharomyces cerevisiae through combining pathway engineering with host engineering, Microbial Cell Factories, vol.8, issue.10, p.113, 2016.
DOI : 10.1371/journal.pone.0075164

S. Chi, Y. He, J. Ren, Q. Su, X. Liu et al., Overexpression of a bifunctional enzyme, CrtS, enhances astaxanthin synthesis through two pathways in Phaffia rhodozyma, Microbial Cell Factories, vol.123, issue.1, p.90, 2015.
DOI : 10.1016/j.jbiotec.2005.11.014

L. Dufossé, Microbial production of food grade pigments, Food Technol. Biotechnol, vol.44, pp.313-321, 2006.

L. Dufossé, Pigments, microbial, Encyclopedia of Microbiology, pp.457-471, 2009.

L. Dufossé and C. De-echanove, The last step in the biosynthesis of aryl carotenoids in the cheese ripening bacteria Brevibacterium linens ATCC 9175 (Brevibacterium aurantiacum sp. nov.) involves a cytochrome P450-dependent monooxygenase, Food Research International, vol.38, issue.8-9, pp.967-973, 2005.
DOI : 10.1016/j.foodres.2005.02.017

L. Dufossé, M. Fouillaud, Y. Caro, S. A. Mapari, and N. Sutthiwong, Filamentous fungi are large-scale producers of pigments and colorants for the food industry, Current Opinion in Biotechnology, vol.26, pp.56-61, 2014.
DOI : 10.1016/j.copbio.2013.09.007

L. Dufossé, P. Galaup, A. Yaron, S. M. Arad, P. Blanc et al., Microorganisms and microalgae as sources of pigments for food use: a scientific oddity or an industrial reality? Trend. Food Sci, pp.389-406, 2005.

N. T. Eriksen, Production of phycocyanin???a pigment with applications in biology, biotechnology, foods and medicine, Applied Microbiology and Biotechnology, vol.8, issue.1, pp.1-14, 2008.
DOI : 10.4319/lo.2005.50.1.0237

J. C. Frisvad, J. Smedsgaard, T. O. Larsen, and R. A. Samson, Mycotoxins, drugs and other extrolites produced by species in Penicillium subgenus Penicillium, Stud. Mycol, vol.49, pp.201-241, 2004.

J. C. Frisvad, N. Yilmaz, U. Thrane, K. B. Rasmussen, J. Houbraken et al., Talaromyces atroroseus, a New Species Efficiently Producing Industrially Relevant Red Pigments, PLoS ONE, vol.22, issue.12, p.84102, 2013.
DOI : 10.1371/journal.pone.0084102.s001

G. Fu, Y. Xu, Y. Li, and W. Tan, Construction of a replacement vector to disrupt pksCT gene for the mycotoxin citrinin biosynthesis in Monascus aurantiacus and maintain food red pigment production. Asia Pac, J. Clin. Nutr, pp.16-137, 2007.

J. M. Gao, S. X. Yang, and J. C. Qin, Azaphilones: Chemistry and Biology, Chemical Reviews, vol.113, issue.7, pp.4755-4811, 2013.
DOI : 10.1021/cr300402y

N. N. Gessler, A. S. Egorova, and T. A. Belozerskaya, Fungal anthraquinones, Applied Biochemistry and Microbiology, vol.49, issue.2, pp.85-99, 2013.
DOI : 10.1134/S000368381302004X

T. L. Grassi, E. F. Do-espírito-santo, M. T. De-siqueira-marcos, J. F. Cavazzana, D. L. Oliveira et al., Bacterial pigment for Nile tilapia feeding, Aquaculture International, vol.10, issue.2, pp.647-660, 2016.
DOI : 10.3923/javaa.2011.1875.1882

A. C. Guedes, H. M. Amaro, and F. X. Malcata, Microalgae as Sources of Carotenoids, Marine Drugs, vol.47, issue.12, pp.625-644, 2011.
DOI : 10.1016/j.cep.2007.07.008

F. Guyomarc-'h, A. Binet, and L. Dufossé, Production of carotenoids by Brevibacterium linens : variation among strains, kinetic aspects and HPLC profiles, Journal of Industrial Microbiology and Biotechnology, vol.24, issue.1, pp.64-70, 2000.
DOI : 10.1038/sj.jim.2900761

S. A. Heider, P. Peters-wendisch, R. Netzer, M. Stafnes, T. Brautaset et al., Production and glucosylation of C50 and C40 carotenoids by metabolically engineered Corynebacterium glutamicum, Applied Microbiology and Biotechnology, vol.17, issue.5, pp.98-1223, 2014.
DOI : 10.1016/j.ymben.2013.02.002

S. A. Heider, P. Peters-wendisch, V. F. Wendisch, J. Beekwilder, and T. Brautaset, Metabolic engineering for the microbial production of carotenoids and related products with a focus on the rare C50 carotenoids, Applied Microbiology and Biotechnology, vol.110, issue.9, pp.4355-4368, 2014.
DOI : 10.1002/bit.24900

I. Jaswir, D. Noviendri, R. F. Hasrini, and F. Octavianti, Carotenoids: sources, medicinal properties and their application in food and nutraceutical industry, J. Med. Plant Res, vol.5, issue.33, pp.7119-7131, 2011.

W. Jin, X. Xu, L. Jiang, Z. Zhang, S. Li et al., Putative carotenoid genes expressed under the regulation of Shine???Dalgarno regions in Escherichia coli for efficient lycopene production, Biotechnology Letters, vol.35, issue.11, pp.37-2303, 2015.
DOI : 10.1007/s10529-013-1317-0

S. H. Kim, M. S. Kim, B. Y. Lee, and P. C. Lee, Generation of structurally novel short carotenoids and study of their biological activity, Scientific Reports, vol.5, issue.1, p.21987, 2016.
DOI : 10.1039/b905598n

M. Kuddus, P. Singh, G. Thomas, and A. Hazimi, Recent Developments in Production and Biotechnological Applications of C-Phycocyanin, BioMed Research International, vol.5, issue.1, p.742859, 2013.
DOI : 10.1155/2013/916590

M. Kumar, P. Dwivedi, A. K. Sharma, M. Sankar, R. D. Patil et al., Apoptosis and lipid peroxidation in ochratoxin A- and citrinin-induced nephrotoxicity in rabbits, Toxicology and Industrial Health, vol.61, issue.2, pp.90-98, 2014.
DOI : 10.1083/jcb.4.4.475

B. M. Lange, T. Rujan, W. Martin, and R. Croteau, Isoprenoid biosynthesis: The evolution of two ancient and distinct pathways across genomes, Proc. Natl. Acad. Sci. USA 97, pp.13172-13177, 2000.
DOI : 10.1046/j.1365-2958.2000.02004.x

X. Li, G. Tian, H. Shen, and J. Liu, Metabolic engineering of Escherichia coli to produce zeaxanthin, Journal of Industrial Microbiology & Biotechnology, vol.29, issue.6, pp.627-636, 2015.
DOI : 10.1016/j.biotechadv.2011.05.020

J. Lin, D. Lee, and J. S. Chang, Lutein production from biomass: Marigold flowers versus microalgae, Bioresource Technology, vol.184, pp.421-428, 2014.
DOI : 10.1016/j.biortech.2014.09.099

I. Mannazzu, S. Landolfo, T. L. Da-silva, and P. Buzzini, Red yeasts and carotenoid production: outlining a future for non-conventional yeasts of biotechnological interest, World Journal of Microbiology and Biotechnology, vol.58, issue.2, pp.31-1665, 2015.
DOI : 10.1590/S1516-8913201400152

S. A. Mapari, A. S. Meyer, U. Thrane, and J. C. Frisvad, Identification of potentially safe promising fungal cell factories for the production of polyketide natural food colorants using chemotaxonomic rationale. Microb. Cell Fact, p.24, 2009.

S. A. Mapari, U. Thrane, and A. S. Meyer, Fungal polyketide azaphilone pigments as future natural food colorants?, Trends in Biotechnology, vol.28, issue.6, pp.300-307, 2010.
DOI : 10.1016/j.tibtech.2010.03.004

L. N. Nupur, A. Vats, S. K. Dhanda, G. P. Raghava, A. K. Pinnaka et al., ProCarDB: a database of bacterial carotenoids, BMC Microbiology, vol.14, issue.10, p.96, 2016.
DOI : 10.1111/j.1462-2920.2012.02806.x

C. Pumas, Y. Peerapornpisal, P. Vacharapiyasophon, P. Leelapornpisid, W. Boonchum et al., Purification and characterization of a thermostable phycoerythrin from hot spring cyanobacterium Leptolyngbya sp, KC45. Int. J. Agric. Biol, vol.14, pp.121-125, 2012.

G. Raguenes, X. Moppert, L. Richert, J. Ratiskol, C. Payri et al., A novel exopolymer-producing bacterium, Paracoccus zeaxanthinifaciens subsp payriae, isolated from a " kopara " mat located in Rangiroa, an atoll of French Polynesia, Curr. Microbiol, vol.49, pp.145-151, 2004.

G. Sandmann, Carotenoids of Biotechnological Importance, Adv. Biochem. Eng. Biotechnol, vol.148, pp.449-467, 2015.
DOI : 10.1007/10_2014_277

E. Sardaryan, H. Zihlova, R. Strnad, and Z. Cermakova, Arpink Red?meet a new natural red food colorant of microbial origin, Pigments in Food, pp.207-208, 2004.

C. Schmidt-dannert, D. Umeno, and F. H. Arnold, Molecular breeding of carotenoid biosynthetic pathways, Nature Biotechnology, vol.18, issue.7, pp.750-753, 2000.
DOI : 10.1038/77319

T. P. Silva, S. M. Paixão, and L. Alves, Ability of Gordonia alkanivorans strain 1B for high added value carotenoids production, RSC Adv., vol.5, issue.63, pp.58055-58063, 2016.
DOI : 10.1039/C4RA14216K

G. H. Song, S. H. Kim, B. H. Choi, S. J. Han, and P. C. Lee, Heterologous Carotenoid-Biosynthetic Enzymes: Functional Complementation and Effects on Carotenoid Profiles in Escherichia coli, Applied and Environmental Microbiology, vol.79, issue.2, pp.610-618, 2013.
DOI : 10.1128/AEM.02556-12

M. Takemura, T. Maoka, A. Osawa, H. Higashinaka, H. Shimada et al., (6E) and (6Z)- 9?-aporhodoxanthinone, novel carotenoids produced in zeaxanthin-synthesizing-Escherichia coli by redox stress, Tetrahedron Lett, issue.44, pp.56-6063, 2015.

P. Thawornwiriyanun, S. Tanasupawat, C. Dechsakulwatana, S. Techkarnjanaruk, and W. Suntornsuk, Identification of Newly Zeaxanthin-Producing Bacteria Isolated from Sponges in the Gulf of Thailand and their Zeaxanthin Production, Applied Biochemistry and Biotechnology, vol.25, issue.8, pp.2357-2368, 2012.
DOI : 10.1023/A:1024199213519

A. Valla, D. Cartier, B. Valla, L. Guillou, R. Andriamialisoa et al., New Synthesis of Natural Carotene Isorenieratene (??,??-Carotene) and its 3,3???-Dimethoxy Analogue, Helvetica Chimica Acta, vol.86, issue.10, pp.3314-3319, 2003.
DOI : 10.1002/hlca.200390273

S. Wagener, T. Völker, S. De-spirt, H. Ernst, and W. Stahl, 3,3???-Dihydroxyisorenieratene and isorenieratene prevent UV-induced DNA damage in human skin fibroblasts, Free Radical Biology and Medicine, vol.53, issue.3, pp.457-463
DOI : 10.1016/j.freeradbiomed.2012.05.022

F. Wang, J. G. Jiang, and Q. Chen, Progress on molecular breeding and metabolic engineering of biosynthesis pathways of C30, C35, C40, C45, C50 carotenoids, Biotechnology Advances, vol.25, issue.3, pp.211-222, 2007.
DOI : 10.1016/j.biotechadv.2006.12.001

P. C. Woo, C. W. Lam, E. W. Tam, K. C. Lee, K. K. Yung et al., The biosynthetic pathway for a thousand-year-old natural food colorant and citrinin in Penicillium marneffei, Scientific Reports, vol.87, issue.1, p.6728, 2014.
DOI : 10.1007/s10482-004-3194-y

Y. Yang, B. Liu, X. Du, P. Li, B. Liang et al., Complete genome sequence and transcriptomics analyses reveal pigment biosynthesis and regulatory mechanisms in an industrial strain, p.8331, 2015.

J. H. Zhang, L. L. Lu, L. J. Yin, S. Xie, and M. Xiao, Carotenogenesis gene cluster and phytoene desaturase catalyzing both three- and four-step desaturations from Rhodobacter azotoformans, FEMS Microbiology Letters, vol.333, issue.2, pp.138-145, 2012.
DOI : 10.1111/j.1574-6968.2012.02604.x

P. Zhou, L. Ye, W. Xie, X. Lv, and H. Yu, Highly efficient biosynthesis of astaxanthin in Saccharomyces cerevisiae by integration and tuning of algal crtZ and bkt, Applied Microbiology and Biotechnology, vol.41, issue.9, pp.99-8419, 2015.
DOI : 10.1093/nar/gkt151

Q. Zhu and E. N. Jackson, Metabolic engineering of Yarrowia lipolytica for industrial applications, Current Opinion in Biotechnology, vol.36, pp.65-72, 2015.
DOI : 10.1016/j.copbio.2015.08.010